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doi:10.2204/iodp.proc.303306.213.2009

Systematics

A list of all Late Quaternary ostracodes found at Site U1314 with key references used for their identification is presented below following the suprageneric taxonomy provided by Whatley and Coles (1987) and updated using the Integrated Taxonomic Information System (www.itis.gov). High-quality SEM photomicrographs are provided for most species. Notes on geographical distribution, ecology, and interpretation can be found in Alvarez Zarikian et al. (2009).

Phylum ARTHROPODA Siebold and Stannius, 1845

Subphylum CRUSTACEA Pennant, 1777

Class OSTRACODA Latreille, 1802

Subclass MYODOCOPA Sars, 1866

Order HALOCYPRIDA Dana, 1853

Suborder CLADOCOPINA Sars, 1866

Family POLYCOPIDAE Sars, 1866

Genus Polycope Sars, 1866

Polycope cf. P. clathrata Joy and Clark (1977), Pl. P1, fig. 6 (Whatley et al., 1998).

Polycope orbicularis Sars, 1866, Pl. P1, fig. 7 (Whatley et al., 1998; Stepanova, 2006).

Polycope punctata Sars, 1870 (Joy and Clark, 1977).

Remarks: Polycope is found during glacial and stadial intervals. The downcore distribution trends are similar to those found in other late Quaternary sequences in the subpolar North Atlantic (Didié and Bauch, 2000, 2002; Didié et al., 2002) where the genus is considerably more abundant (Alvarez Zarikian et al., 2009).

Order PODOCOPIDA Müller, 1894

Suborder CYTHEROCOPINA Sars, 1866

Superfamily CYTHEROIDEA Baird, 1850

Family BYTHOCYTHERIDAE Sars, 1866

Genus Bythocythere Sars, 1866

Bythocythere bathytatos Whatley and Coles, 1987, Pl. P2, fig. 3 (Whatley and Coles, 1987).

Remarks: B. bathytatos is most abundant during marine isotope stages (MIS) 6, 5d, and 2. This species was first recovered from upper Quaternary sediments at >3400 m water depth in the central North Atlantic (Whatley and Coles, 1987; Cronin et al., 1999). It was recently reported from very cold waters (1.2°C) 5032 m east of the Walvis Ridge in the eastern South Atlantic (Yasuhara et al., 2008b). B. bathytatos also shows a glacial/stadial distribution in the upper Pleistocene sediments at the Rockall Plateau in the eastern North Atlantic (Didié and Bauch, 2000).

Genus Pseudocythere Sars, 1866

Pseudocythere caudata Sars, 1866, Pl. P2, fig. 4 (Whatley et al., 1998; Stepanova, 2006).

Pseudocythere sp. 2 sensu Didié and Bauch (2000), Pl. P2, fig. 6.

Remarks: Species of Pseudocythere were found in samples corresponding to glacial intervals. P. caudata has been found worldwide in the deep ocean and it is common in the Arctic Ocean (Joy and Clark, 1977; Didié and Bauch, 2000). Pseudocythere sp. 2 has also been found in glacial sediments from the Rockall Plateau (Didié and Bauch, 2000).

Family CYTHERIDAE Baird, 1850

Genus Nannocythere Schäfer, 1953

Nannocythere sp., Pl. P2, fig. 1 (Whatley et al., 1998; Didié and Bauch, 2000, 2001).

Remarks: Nannocythere sp. is found in very low numbers during interglacials.

Family CYTHERIDEIDAE Sars, 1925

Genus Heterocyprideis Elofson, 1941

Heterocyprideis sorbyana (Jones) 1857 (Cronin, 1981; Stepanova, 2006).

Remarks: A single valve of H. sorbyana was found in sediments corresponding to the last glacial interval (MIS 2). This species lives in cold regions of the Atlantic and in the Arctic Ocean (Cronin, 1981).

Family CYTHERURIDAE Müller, 1894

Genus Aversovalva Hornibrook, 1952

Aversovalva hydrodynamica Whatley and Coles, 1987, Pl. P3, fig. 7 (Whatley and Coles, 1987).

Remarks: One complete carapace was found in a sample corresponding to the Holocene. The species was also found in interglacial sediments at the Rockall Plateau (Didié and Bauch, 2000).

Genus Cytheropteron Sars, 1866

Cytheropteron alatum Sars, 1866, Pl. P3, fig. 4 (Joy and Clark, 1977; Didié and Bauch, 2000, 2001).

Cytheropteron arcuatum Brady, Crosskey, and Robertson, 1874, Pl. P4, fig. 8 (Whatley and Masson, 1979; Stepanova et al., 2004).

Cytheropteron carolinae Whatley and Coles, 1987, Pl. P4, fig. 7 (Whatley and Coles, 1987).

Cytheropteron champlainum Cronin, 1981 (Cronin, 1981).

Cytheropteron circummuralla Whatley and Coles, 1987, Pl. P4, figs. 3, 4 (Whatley and Coles, 1987).

Cytheropteron hamatum Sars, 1869 (Whatley et al., 1998).

Cytheropteron lineoporosa Whatley and Coles, 1987, Pl. P3, figs. 3, 5 (Whatley and Coles, 1987).

Cytheropteron massoni Whatley and Coles, 1987 (Whatley and Coles, 1987).

Cytheropteron perlaria Hao, 1988, Pl. P3, figs. 1, 2 (Hao, 1988; Stepanova, 2006; as C. testudo in Whatley and Coles, 1987).

Cytheropteron pherozigzag Whatley and Ayress, 1988 (Whatley and Coles, 1987).

Cytheropteron porterae Whatley and Coles, 1987, Pl. P4, figs. 5, 6 (Whatley and Coles, 1987).

Cytheropteron syntomalatum Whatley and Coles, 1987, Pl. P3, fig. 6 (Whatley and Coles, 1987).

Cytheropteron tenuialatum Whatley and Coles, 1987, Pl. P4, figs. 1, 2 (Whatley and Coles, 1987).

Cytheropteron tressleri Whatley and Coles, 1987 (Whatley and Coles, 1987).

Cytheropteron trifossata Whatley and Coles, 1987 (Whatley and Coles, 1987).

Remarks: More than 14 species of Cytheropteron were found in late Quaternary sediments at Site U1314, making it the most diverse genus at the site. Most common species are C. syntomalatum, C. porterae, C. perlaria, C. tenuialatum, and C. circummuralla. Although found throughout the entire stratigraphic section, Cytheropteron species are most abundant during MIS 6, 4, and 2, particularly during deglaciations (Alvarez Zarikian et al., 2009). Lowest abundances (<5%) are observed during the Eemian, the Holocene, and early in MIS 3. C. arcuatum, C. champlainum, C. lineoporosa, and C. trifossata are found as single specimens in samples with high IRD content and associated with deglaciations. C. arcuatum and C. champlainum are commonly found at shallower depths on the shelf/upper slope in the North Atlantic and Arctic seas (Stepanova et al., 2004, and references therein).

Genus Eucytherura Müller, 1894

Eucytherura calabra (Cotalongo and Pasini) 1980 (Whatley and Coles, 1987).

Remarks: Widespread in deep-sea sediments worldwide, it was found as single specimens in only four samples.

Genus Pedicythere Eagar, 1965

Pedicythere polita Colalongo and Pasini, 1980 (Whatley and Coles, 1987).

Remarks: Members of this genus were previously reported from deep-sea sediments in the deep North Atlantic (Cronin, 1983; Whatley and Coles, 1987), Greenland Sea (Whatley et al., 1998), and Arctic Ocean (Joy and Clark, 1977; Jones et al., 1998). Pedicythere polita was found in low numbers in samples corresponding to glacial and stadial intervals.

Genus Pelecocythere Athersuch, 1979

Pelecocythere sylvesterbradleyi Athersuch, 1979, Pl. P5, fig. 4 (Athersuch, 1979; Whatley and Coles, 1987).

Remarks: Pelecocythere sylvesterbradleyi is a common deep-water species in the North Atlantic (Neale, 1988). It exhibited maximum abundances during MIS 5e at Site U1314 and was reported as the major component of the “interglacial” assemblage in upper Quaternary sediments in the Iceland Plateau (Didié et al., 2002). Similarly, it has also been reported as a typical component of interglacial assemblages at other North Atlantic sites (e.g., Cronin et al., 1996, 1999; Didié and Bauch, 2000).

Genus Rimacytheropteron Whatley and Coles, 1987

Rimacytheropteron longipunctata (Breman) 1976. Pl. P3, fig. 8 (Whatley and Coles, 1987).

Remarks: A single specimen of R. longipunctata was found in a core sample corresponding to ~145 ka.

Genus Semicytherura Wagner, 1957

Remarks: A single valve of an unidentified species of Semicytherura was found in a core sample corresponding to ~70 ka.

Genus Swainocythere Ishizaki, 1981

Swainocythere nanseni (Joy and Clark) 1981, Pl. P2, fig. 7 (as Cytheroteron? nealei in Joy and Clark, 1977; Corrège et al., 1992; Didié and Bauch, 2000, 2001).

Swainocythere sp.1 sensu Didié and Bauch (2000, 2001) Pl. P2, fig. 8.

Remarks: Species of Swainocythere were found in low numbers in ~10 samples. S. nanseni is found today in the deep Arctic Ocean (Joy and Clark, 1977; Jones et al., 1998) and around Australia (Corrège et al., 1992).

Family EUCYTHERIDAE Puri, 1954

Genus Eucythere Brady, 1868

Eucythere argus (Sars) 1866 (Cronin, 1981; Whatley et al., 1998).

Eucythere circumcostata Whatley and Coles, 1987, Pl. P6, fig. 2 (Whatley and Coles, 1987).

Eucythere multipunctata Whatley and Coles, 1987, Pl. P6, fig. 1 (Whatley and Coles, 1987).

Eucythere pubera Bonaduce, Ciampo, and Masoli, 1975, Pl. P6, fig. 3 (Whatley and Coles, 1987).

Eucythere triangula Whatley and Coles, 1987, Pl. P6, fig. 4 (Whatley and Coles, 1987).

Eucythere sp. B sensu Whatley et al. (1998), Pl. P6, fig. 5.

Remarks: Species of Eucythere occur in only a few samples associated with elevated IRD content, where they reached up to 7% of the assemblage. These intervals correspond to MIS 5d, 5b, 4, and 2 (Alvarez Zarikian et al., 2009). Species of Eucythere have been recovered from water depths from 650 to >4000 m from the Greenland Sea (Whatley et al., 1998) to the middle North Atlantic (Whatley and Coles, 1987) and many of them are restricted at depths >3000 m (Dingle and Lord, 1990). Eucythere has been recorded at glacial–interglacial transitions in upper Quaternary sediment sequences from the polar and subpolar North Atlantic (Didié and Bauch, 2000; Didié et al., 2002).

Family HEMICYTHERIDAE Puri, 1953

Genus Finmarchinella Swain, 1963

Finmarchinella finmarchica (Sars) 1865, Pl. P6, figs. 6, 7 (Cronin, 1991).

Remarks: This species was found as single valves in samples with high IRD content. Members of this genus are very abundant in the Northern Seas (Neale and Howe, 1975; Cronin, 1991; Didié and Bauch, 2000).

Genus Hemicythere Sars, 1925

Hemicythere villosa (Sars) 1866, Pl. P6, fig. 8 (Athersuch et al., 1989; Didié and Bauch, 2001).

Remarks: A single valve of H. villosa was found in a sample corresponding to ~41 ka. Amphiatlantic species are found along the Arctic coasts (Cronin et al., 1993).

Family KRITHIDAE Mandelstam, 1958

Genus Krithe Brady, Crosskey, and Robertson, 1874

Krithe cf. K. aequabilis Campo, 1986, Pl. P7, fig. 2 (Coles et al., 1994; Rodriguez-Lázaro et al., 1999).

Krithe cf. K. dolichodeira van den Bold, 1946, Pl. P7, fig. 3, 4 (Coles et al., 1994) .

Krithe cf. K. minima Coles, Whatley, and Moguilevsky, 1994 (Coles et al., 1994; Rodriguez-Lázaro et al., 1999).

Krithe morkhoveni van den Bold, 1960, Pl. P7, fig. 1 (Coles et al., 1994; Rodriguez-Lázaro et al., 1999).

Krithe trinidadensis van den Bold, 1958, Pl. P7, fig. 5 (Coles et al., 1994; Rodriguez-Lázaro et al., 1999).

Remarks: The genus Krithe in our samples was represented by at least 3–4 species. It was most dominant during the 150–165 ka interval in MIS 6, the early half of MIS 5e, immediately after peak stadial (MIS 5d and 5b), during most of MIS 3, and the early part of the Holocene (Alvarez Zarikian et al., 2009).

Family LOXOCONCHIDAE Sars, 1925

Genus Heinia van den Bold, 1985

Heinia dryppa Whatley and Coles, 1987, Pl. P2, fig. 2 (Whatley and Coles, 1987).

Remarks: Few specimens, including one complete carapace, were found in a core sample dated at 120 ka. This species also occurs in the southwest Pacific and Indian Ocean (Whatley and Coles, 1987).

Genus Loxoconcha Sars, 1866

?Loxoconcha sp., Pl. P7, figs. 7, 8.

Remarks: An unidentified species resembling a Loxoconchiid occurs in several core samples corresponding to the Holocene and the last interglacial.

Family MICROCYTHERIDAE Klie, 1938

Genus Microcythere Müller, 1894

Microcythere medistriata (Joy and Clark, 1977), Pl. P8, figs. 7, 8 (Joy and Clark, 1977).

Remarks: A single carapace of M. medistriata was found in a sample corresponding to ~8 ka.

Family PARADOXOSTOMATIDAE Brady and Norman, 1889

Genus Cytherois Müller, 1884

Cytherois pusilla Sars 1928.

Remarks: Three valves of this species were found in the core section in intervals corresponding to deglaciations.

Genus Paracytherois Müller, 1894

Paracytherois flexuosa (Brady) 1867, Pl. P8, fig. 9 (Whatley et al., 1998).

Remarks: Few specimens of this species were found in the core section. P. flexuosa has been previously reported from the Greenland Sea shelf (Whatley et al., 1998).

Family ROCKALLIIDAE Whatley et al., 1982

Genus Rockallia Whatley, Frame, and Whittaker, 1978

Rockallia enigmatica Whatley, Frame, and Whittaker, 1978, Pl. P9, fig. 5 (Whatley and Coles, 1987; Whatley et al., 1982; Ayress, 1991).

Remarks: R. enigmatica is one of the most abundant species at Site U1314, reaching >20% of the total assemblage during MIS 3 and glacial and deglacial transitions. In contrast, it was nearly absent during full interglacial and glacial intervals (Alvarez Zarikian et al., 2009). R. enigmatica shows a similar stratigraphic distribution on the southern Rockall Plateau but it was generally less abundant there (Didié and Bauch, 2000). It is considered a bathyal and abyssal species found in the North Atlantic (Whatley et al., 1982). The genus Rockallia is a junior synonym of the genus Arcacythere Hornibrook, 1952 (Ayress, 1991), but I kept Rockallia in this report for consistency with the North Atlantic literature.

Family TRACHYLEBERIDIDAE Sylvester-Bradley, 1948

Genus Ambocythere van den Bold, 1957

Ambocythere ramosa van den Bold, 1965, Pl. P10, fig. 7 (Whatley and Coles, 1987).

Remarks: A. ramosa reached highest proportions during MIS 5e and 5c and the Holocene (Alvarez Zarikian et al., 2009). It was common in deep-sea Core M23414 from the southern Rockall Plateau, where it reached its greatest relative abundance (3.5%) during MIS 5 (Didié and Bauch, 2000). A. ramosa was also reported from the North Atlantic deep waters (Whatley and Coles, 1987).

Genus Bathycythere Sissingh 1971

Bathycythere audax (Brady and Norman, 1869; as B. vanstraateni Sissingh 1971 in Dingle and Lord, 1990), Pl. P5, fig. 5 (Whatley and Coles, 1987).

Remarks: B. audax peaked in core samples corresponding to MIS 6, 4, and 2 (Alvarez Zarikian et al., 2009). This species is restricted to water depths >3000 m in the Atlantic and temperatures <3°C. Its upper depth distribution limits are restricted to the lowermost part of the NADW (Dingle and Lord, 1990). In the southwest Pacific and South Indian oceans, B. audax has been found between 1900 and 3584 m in coldwater masses associated with the Deep Water and Antarctic Bottom Water (AABW) (Ayress et al., 2004).

Genus Bradleya Hornibrook, 1952

Bradleya dictyon (Brady) 1880, Pl. P5, figs. 8, 9; Pl. P9, fig. 1 (Whatley and Coles, 1987).

Bradleya normani Brady, 1865, Pl. P9, fig. 2 (Benson, 1972).

Remarks: Two species of Bradleya were recognized. The genus displays an interglacial and interstadial distribution pattern, reaching maximum relative abundances (~15%) during MIS 5e (Alvarez Zarikian et al., 2009). The species are known from the deep waters of the Atlantic Ocean (Guernet and Fourcade, 1988).

Genus Dutoitella Dingle, 1981

Dutoitella suhmi (Brady, 1880), Pl. P5, figs. 6, 7 (Whatley and Coles, 1987).

Remarks: Abyssal species D. suhmi peaked in discrete samples during MIS 6, 4, and 2 (Alvarez Zarikian et al., 2009). In the Atlantic, this species is restricted to water depths >3000 m and temperatures <3°C. Its upper depth limits are restricted to the lowermost part of the NADW (Dingle and Lord, 1990). In the southwest Pacific and South Indian oceans, D. suhmi has been reported from a water depth >3584 m and temperatures <1°C (Ayress et al., 2004).

Genus Echinocythereis Puri, 1954

Echinocythereis echinata (Sars) 1866, Pl. P9, figs. 3, 4 (Whatley and Coles, 1987).

Remarks: E. echinata reached highest relative abundances (>7%) during an interval in MIS 6, as well as during MIS 4. However, it was also present, though slightly less abundant, during MIS 5e, 5c, and 3 (Alvarez Zarikian et al., 2009). E. echinata appears to be restricted to the region between mid and low latitudes in the Atlantic at water depths ranging from 500 to 3884 m (Dingle and Lord, 1990; Didié et al., 2002). This species has been associated with interglacial assemblages in the central (Cronin et al., 1999) and northeastern North Atlantic (Didié and Bauch, 2000).

Genus Henryhowella Puri, 1957

Henryhowella asperrima (Reuss) 1850 (Whatley and Coles, 1987).

Henryhowella dasyderma (Brady) 1880, Pl. P9, figs. 6, 7, 8 (Whatley and Coles, 1987).

Remarks: Genus Henryhowella (mainly H. cf. dasyderma with occasionally occurring H. cf. asperrima) was consistently present throughout the cored section (average ~5%) reaching maximum relative abundances (>15%) during MIS 5e, 5a, 3, and the Holocene (Alvarez Zarikian et al., 2009).

Genus Legitimocythere Coles and Whatley, 1989 (= Genus “Thalassocythere” Benson, 1977)

Legitimocythere acanthoderma (Brady) 1880, Pl. P1, figs. 4, 5 (Whatley and Coles, 1987).

Remarks: Legitimocythere acanthoderma was frequently found in relatively lower abundance (<5%). Its occurrence appears to be associated with glacials and stadials (Alvarez Zarikian et al., 2009). This species is reported as “Thalassocythere” acanthoderma in Coles and Whatley, 1989 (Dingle and Lord, 1990).

Genus Pennyella Neale, 1974 (= Genus “Oxycythereis” Benson, 1974)

Pennyella dorsoserrata (Brady) 1880, Pl. P1, figs. 1, 2 (Whatley and Coles, 1987).

Pennyella horridus Whatley and Coles, 1987, Pl. P1, fig. 3 (Whatley and Coles, 1987).

Remarks: Genus Pennyella (as Oxycythereis in Whatley and Coles, 1987; Cronin et al., 1999; Didié and Bauch, 2000; and as Rugocythereis in Dingle and Lord, 1990) is represented by P. dorsoserrata and P. horridus. This genus was present in sediments deposited during MIS 5a, 5c, 5e, and the Holocene and clearly showed its preference to interstadial and interglacial conditions (Alvarez Zarikian et al., 2009). It was also abundant in upper Pleistocene to Holocene sediments on the southern Rockall Plateau (Didié and Bauch, 2000).

Genus Pseudobosquetina Guernet and Moullade, 1994

Pseudobosquetina mucronalatum (Brady) 1880 (Whatley and Coles, 1987; Jellinek et al., 2006).

Pseudobosquetina nobilis Jellinek, Swanson, and Mazzini, 2006, Pl. P5, figs. 1, 2, 3 (Jellinek et al., 2006).

Remarks: Specimens of genus Pseudobosquetina found at Site U1314 were identified as P. mucronalatum in Alvarez Zarikian et al. (2009) (as Bosquetina mucronalatum in Whatley and Coles, 1987; Didié and Bauch, 2000, 2001; as Pterygocythere mucronalatum in Ayress et al., 2004), but further examination showed that some of the specimens may also belong to P. nobilis Jellinek, Swanson, and Mazzini, 2006. Pseudobosquetina showed highest abundances (>12%) during MIS 5e and occurred sporadically at lower levels during MIS 5b–5d, 6, 4, 3, 2, and 1 (Alvarez Zarikian et al., 2009). Pseudobosquetina is a cosmopolitan abyssal genus commonly found in the Atlantic and Pacific oceans. It is considered to be an important component of ostracode assemblages found in sediments bathed by lower NADW and AABW for its modern bathymetric distribution (2400 to >4600 m) and association with deepwater masses (Dingle and Lord, 1990; Ayress et al., 1997; Yasuhara et al., 2008b). P. nobilis occurs in the North Atlantic and the southeast Atlantic (Jellinek et al., 2006).

Family XESTOLEBERIDAE Sars, 1928

Genus Xestoleberis Sars, 1866

Xestoleberis profundis Whatley and Coles, 1987, Pl. P7, fig. 6 (Whatley and Coles, 1987).

Remarks: Xestoleberis profundis peaked at the MIS 3–2 and MIS 6–5 transitions. It has been recovered from the central North Atlantic (Whatley and Coles, 1987; Van Harten, 1990) and South Atlantic (Yasuhara et al., 2008b) at depths ranging from 2600 to 5000 m.

Suborder Bairdiocopina

Superfamily BAIRDIOIDEA Sars, 1887

Family BAIRDIIDAE Sars, 1887

Genus Bairdoppilata Coryell, Sample, and Jennings, 1935

Bairdoppilata victrix (Brady) 1954, Pl. P10, figs. 1, 2 (Whatley and Coles, 1987).

Remarks: Bairdoppilata victrix was found in core samples associated with glacial and stadial intervals. This species is found in deep waters in the North Atlantic (Whatley and Coles, 1987; Didié, 2001) and has been linked to glacial assemblages on the Rockall Plateau (Didié and Bauch, 2000).

Family BYTHOCYPRIDIDAE Maddocks, 1969

Genus Bythocypris Brady, 1880

(Pl. P10, figs. 4, 5, 6)

Remarks: Specimens of this genus were found sporadically throughout the core section in intervals typically associated with glacial stages. This is a widely distributed deep-sea ostracode.

Suborder Cypridocopina Jones, 1901 Superfamily Macrocypridoidea Müller, 1912

Family MACROCYPRIDIDAE Müller, 1912

Genus Macrocypris Brady, 1868

(Whatley and Coles, 1987)

Remarks: Scattered findings in low numbers.

Superfamily Pontocypridoidea Müller, 1894

Family PONTOCYPRIDIDAE Müller, 1894

Genus Argilloecia Sars, 1866

Argilloecia spp., Pl. P8, figs. 1–6 (Whatley and Coles, 1987).

Remarks: At least four different species of Argilloecia were recognized (three are illustrated in Pl. P8). Two correspond to Argilloecia sp. 5 and Argilloecia sp. 6 distinguished by Whatley and Coles (1987) in sediments cored at Deep Sea Drilling Project Leg 94 sites in the North Atlantic. Highest relative abundances of this genus are restricted to MIS 5e and 5c (up to 20% of the fauna), while it exhibits slightly lower abundances during Interstadial 5a, the early part of the MIS 3, and the Holocene. Intervals when Argilloecia displays maximum abundances are concurrent with intervals of maximum CaCO₃ and minimum IRD content in the sediments (Alvarez Zarikian et al., 2009).

Genus Propontocypris Sylvester-Bradley, 1947

Propontocypris trigonella (Sars), Pl. P8, fig. 10 (Whatley and Coles, 1987).

Remarks: P. trigonella showed a similar downcore distribution as Argilloecia, but also peaked during an interval in MIS 6 centered at ~145 ka. Most species of Propontocypris are quick swimmers, known to feed on decaying animal and plant tissue (Maddocks and Steineck, 1987), and some of them have been reported from deepwater vents associated with cold-sea biota (personal observation).

Suborder PLATYCOPINA Sylvester-Bradley, 1961

Superfamily CYTHERELLOIDEA Sars, 1866

Family CYTHERELLIDAE Sars, 1866

Genus Cytherella Jones, 1849

Remarks: Two specimens of Cytherella were found throughout the entire stratigraphic section studied. They belong to the same species illustrated in Didié and Bauch (2000, 2001) as Cytherella sp. 1 (Pl. P10, fig. 5).

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