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This research used samples provided by the Integrated Ocean Drilling Program (IODP). We thank the captain, crew, and technicians of the R/V JOIDES Resolution, Co-Chief scientists Katrina J. Edwards and Wolfgang Bach, and the IODP Expedition 336 shipboard scientific party, especially the members of the microbiology team. We also thank Kentaro Nakamura for helpful discussion and Takuro Nunoura and Miho Hirai for their assistance in the mcrA detection procedure.