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We thank the captain and crew of the R/V JOIDES Resolution and the Expedition 327 Shipboard Scientific Party. We also thank Karen Lloyd for providing a copy of her MCG ARB database, Jennifer Biddle and Amanda Martino for their thoughtful reviews, and Beth Orcutt and Geoff Wheat for useful discussion. This research was supported by funding from a Schlanger Ocean Drilling Fellowship to S.P. Jungbluth and an Expedition 327 participation grant (T327A4) to J.P. Cowen, both of which are part of the National Science Foundation (NSF)-sponsored United States Science Support Program for IODP administered by the Consortium for Ocean Leadership, a Center for Dark Energy Biosphere Investigations (C-DEBI; NSF award number 093956-4) Graduate Fellowship to S.P. Jungbluth and research award to M.S. Rappé, and the Center for Microbial Oceanography: Research and Education (C-MORE; award number EF0424599). This study used samples and/or data provided by the Integrated Ocean Drilling Program (IODP). This is SOEST contribution 9024, HIMB contribution 1571, and Center for Dark Energy Biosphere Investigations (C-DEBI) contribution 181.